Bold denotes lab membership at time of research.
* indicates undergraduate mentee.


Wieczynski, D, HV Moeller, and JP Gibert. Mixotrophic microbes create carbon tipping points under warming. Functional Ecology (accepted).

Kaare-Rasmussen, JO*HV Moeller, and F Pfab. Modeling Food Dependent Symbiosis in Exaiptasia pallida. Ecological Modelling 481: 110325. [online open access]

Chu, T*HV Moeller, and KM Archibald. Competition between phytoplankton and mixotrophs leads to metabolic character displacement. Ecological Modelling 481: 110331. [online open access]

Kopecky, KL, AC Stier, RJ Schmitt, SJ Holbrook, and HV Moeller. Material legacies can degrade resilience: Structure-retaining disturbances promote regime shifts on coral reefs. Ecology: e4006. [online open access]

Johnson, MD, HV MoellerC Paight, RM Kellogg, MR McIlvin, MA Saito, and E Lasek-Nesselquist. Functional control and metabolic integration of stolen organelles in a photosynthetic ciliate. Current Biology (in press). [online access] [pdf]

Moeller, HV, RM Nisbet, and AC Stier. Cascading benefits of mutualists' predators on foundation species: a model inspired by coral reef ecosystems. Ecosphere 14 (1): e4382. [online open access] [data & code]


Gonzalez, LM*, SR Proulx, and HV Moeller. Modeling the metabolic evolution of mixotrophic phytoplankton in response to rising ocean surface temperatures. BMC Ecology and Evolution 22: 136. [online open access] [data & code]

Archibald, KM, S Dutkiewicz, C Laufkötter, and HV Moeller. Thermal responses in global marine planktonic food webs are mediated by temperature effects on metabolism. JGR Oceans 127(12): e2022JC018932. [online open access] [data & code]

Lepori-Bui, MC PaightE EberhardCM Mertz*, and HV Moeller. Evidence for evolutionary adaptation of mixotrophic nanoflagellates to warmer temperatures. Global Change Biology 28: 7094-7107. [press release] [online open access] [pdf] [data & code]

Paight, C, MD Johnson, E Lasek-Nesselquist, and HV Moeller. Cascading effects of prey identity on gene expression in a kleptoplastidic ciliate. Journal of Eukaryotic Microbiology 70(1): e12940. [online access]

Pfab, FAL BrownAR DetmerEC Baxter*HV Moeller, R Cunning, and RM Nisbet. Timescale separation and models of symbiosis: state space reduction, multiple attractors, and initialization. Conservation Physiology 10 (1): coac026. [pdf] [online access]

Brown, AL, F Pfab, EC Baxter, AR Detmer, HV Moeller, RM Nisbet, and R Cunning. "Analysis of a mechanistic model of corals in association with multiple symbionts: Within-host competition and recovery from bleaching." Conservation Physiology 10 (1): coac066. [online access]

Hsu, V*F Pfab, and HV Moeller. Niche expansion via acquired metabolism facilitates competitive dominance in planktonic communities. Ecology 103 (7): e3693. [pdf] [press release] [data & code] [online access]

Detmer, AR, R Cunning, F PfabAL Brown, AC Stier, RM Nisbet, and HV Moeller. Fertilization by coral-dwelling fish promotes coral growth but can exacerbate bleaching response. Journal of Theoretical Biology 541: 111087. [pdf] [data & code] [online access]

Falcó, C* and HV Moeller. Optimal spatial management in a multi-use marine habitat: Balancing fisheries and tourism. Natural Resource Modeling 35(1): e12309. [pdf] [online access]


Runte, GCAH Smith*HV Moeller, and LM Bogar. Spheres of influence: Host tree proximity and soil chemistry shape rRNA, but not DNA, communities of symbiotic and free-living soil fungi in a mixed hardwood-conifer forest. Frontiers in Ecology and Evolution 9: 397. [pdf] [online access]

Moeller, HVV Hsu*M Lepori-BuiLY MesropC Chinn*, and MD Johnson. Prey type constrains growth and photosynthetic capacity of the kleptoplastidic ciliate Mesodinium chamaeleon (Ciliophora). Journal of Phycology 57(3): 916-930. [pdf] [data & code] [online access]

Hsu, V*, and HV Moeller. Metabolic symbiosis facilitates species coexistence and generates light-dependent priority effects. Frontiers in Ecology and Evolution 9: 491. [pdf] [online access] [data & code]

Detmer, AR*, RJ Miller, DC Reed, TW Bell, AC Stier, and HV Moeller. Variation in disturbance to a foundation species structures the dynamics of a benthic reef community. Ecology 102(5): e03304. [pdf] [online access] [data & code] [press release]


Bui, A, D Orr, M Lepori-Bui, K Konicek, HS Young, and HV Moeller. Soil fungal community composition and functional similarity shift across distinct climatic conditions. FEMS Microbiology Ecology 96(12): fiaa193. [press release] [pdf] [online access] [data & code]

Pisapia, C, PJ Edmunds, HV Moeller, B Riegl, M McWilliam, CD Wells, and MS Pratchett. Projected shifts in coral size structure in the Anthropocene. Advances in Marine Biology (in press). [online access]

Rice, MM, BL Maher, AMS Correa, HV Moeller, NP Lemoine, AA Shantz, DE Burkepile, and NJ Silbiger. Macroborer presence on corals increases with nutrient input and promotes parrotfish bioerosion. Coral Reefs 39: 409-418. [online access]

Visalli, ME, BD Best, RB Cabral, ... HV Moeller, ... DJ McCauley. Data-driven approach for highlighting priority areas for protection in marine areas beyond national jurisdiction. Marine Policy 122: 103927. [online access]


Moeller, HV, MG Neubert, and MD Johnson. Intraguild predation enables coexistence of competing phytoplankton in a well-mixed water column. Ecology 100 (12): e02874. [pdf] [online access]

Moeller, HV, C Laufkötter, EM Sweeney, and MD Johnson. Light-dependent grazing can drive formation and deepening of Deep Chlorophyll Maxima. Nature Communications 10: 1978. [press release] [pdf]


Pruitt, JN, A Berdahl, C Riehl, N Pinter-Wollman, HV Moeller, et al. Social tipping points in animal societies. Proc. R. Soc. B. 285: 20181282. [pdf]

Johnson, MD and HV Moeller. Mixotrophy in protists: From model systems to mathematical models. Frontiers in Marine Science 5: 490.

Doering, GN, I Scharf, HV Moeller, and JN Pruitt. Social tipping points in animal societies in response to heat stress. Nature Ecology & Evolution 2(8): 1298. [online access]

Moeller, HV and MD Johnson. Preferential plastid retention by the acquired phototroph Mesodinium chamaeleonJournal of Eukaryotic Microbiology 65(2): 148-158. [pdf]


Johnson, MD, E Lasek-Nesselquist, HV Moeller, et al. Mesodinium rubrum: The symbiosis that wasn't. PNAS 114(7): E1040-E1042. [online access]


Moeller, HV and KG Peay. Competition-function tradeoffs in ectomycorrhizal fungi. PeerJ 4:e2270. [online access] [pdf]

Moeller, HV, I Dickie, T Fukami, and D Peltzer. Hierarchical neighbor effects on mycorrhizal community structure and function. Ecology & Evolution 6(15): 5416-5430. [online access] [pdf]

Herrera, G, HV Moeller, and MG Neubert. High-seas fish wars generate marine reservesPNAS 113(14): 3767-3772.

Pedersen, E, J Marleau, M Granados, HV Moeller, F Guichard. Non-heirarchical movement promotes stability and resilience in a tritrophic system. The American Naturalist 187(5): E116-E128.

Moeller, HV, E Peltomaa, MD Johnson, and MG Neubert. Acquired phototrophy stabilizes coexistence and shapes dynamics of an intraguild predator and its prey. Ecology Letters 19: 393-402[pdf]

Moeller, HV, and MG Neubert. Multiple friends with benefits: An optimal mutualist management strategy? The American Naturalist 187(1): E1-E12. [online access] [pdf] [appendices]


Moeller, HV, I Dickie, T Fukami, and D Peltzer. Mycorrhizal co-invasion and novel interactions depend on neighborhood context. Ecology 96(9): 2336-2347[pdf]

Moeller, HV, and MG Neubert. Economically optimal marine reserves without spatial heterogeneity in a simple two-patch model. Natural Resource Modeling 28(3): 244-255. [pdf]

Wood, J, I Dickie, HV Moeller, D Peltzer, K Boot, G Rattray, and J Wilmshurst. Novel interactions between non-native mammals and fungi facilitate establishment of invasive pines. Journal of Ecology 103: 121-129. [pdf]


Moeller, HV, KG Peay, T Fukami. Ectomycorrhizal fungal traits reflect environmental conditions along a coastal California edaphic gradient. FEMS Microbiology Ecology 87(3): 797-806. [pdf]

Listen to a Goggles Optional podcast (complete with fungus song) about this paper here.


Moeller, HV and MG Neubert. Habitat damage, marine reserves, and the value of spatial management. Ecological Applications 23, 959-971. [pdf]

Karp, DS, HV Moeller, and LO Frishkoff. Non-random extinction patterns can modulate pest-control service decline. Ecological Applications 23: 840-849. [pdf]

Sailley, SF, HW Ducklow, HV Moeller, WR Fraser, OME Schofield, DK Steinberg, LM Garzio, and SC Doney. Carbon fluxes and pelagic ecosystem dynamics near two western Antarctic Peninsula Adelie penguin colonies: an inverse model approach. Marine Ecology Progress Series 492: 253-272.


Ducklow, H, SC Doney, ... HV Moeller, et al. Marine Pelagic Ecosystems: The West Antarctic Peninsula. In: Antarctic Ecosystems: An extreme environment in a changing world, 121-159.


Moeller, HV, MD Johnson, PG Falkowski. Photoacclimation in the phototrophic marine ciliate, Mesodinium rubrum (Ciliophora). Journal of Phycology 47: 324-332. [pdf]


Johnson, MD, J Volker, HV Moeller, E Laws, KJ Breslauer, PG Falkowski. Universal constant for heat production in protists. PNAS 106: 6696-6699.